Information about Water Mold

Water molds

Scientific classification
Domain:Eukaryota
Kingdom:Chromalveolata
Phylum:Heterokontophyta
Class:Oomycetes
Orders


Lagenidiales
Leptomitales
Peronosporales
Rhipidiales
Saprolegniales
Sclerosporales
Enlarge picture
A water mold from a stream
Water molds (or water moulds: see spelling differences) also known as Oomycetes are a group of filamentous, unicellular protists, physically resembling fungi. They are microscopic, absorptive organisms that reproduce both sexually and asexually and are composed of mycelia, or a tube-like vegetative body (all of an organism's mycelia are called its thallus). The name "water mold" refers to their earlier classification as fungi, which stemmed from their preference for conditions of high humidity and running surface water, although they are now known to have evolved separately and show a number of differences. For instance, their cell walls are composed of cellulose rather than chitin and generally do not have septations. Also, in the vegetative state they have diploid nuclei, whereas fungi have haploid nuclei.

Instead, water molds are related to organisms such as brown algae and diatoms, making up a group called the heterokonts. The name comes from the common arrangement and structure of motile cells, which typically have two unequal flagella. Among the water molds, these are produced as asexual spores called zoospores, which capitalize on surface water (including precipitation on plant surfaces) for movement. They also produce sexual spores, called oospores, that are translucent double-walled spherical structures used to survive adverse environmental conditions. A few produce aerial asexual spores that are distributed by wind.

The water molds are economically and scientifically important because they are aggressive plant pathogens (see plant pathology). The majority can be broken down into three groups, although more exist.
  • The Phytophthora group is a genus that causes diseases such as dieback, potato blight, sudden oak death, rhododendron root rot, and Ink Disease in the American Chestnut.
  • The Pythium group is even more prevalent than Phytophythora and individual species have larger host ranges, usually causing less damage. Pythium damping off is a very common problem in greenhouses where the organism kills newly emerged seedlings. Mycoparasitic members of this group (e.g. P. oligandrum) parasitize other oomycetes and fungi, and have been employed as biocontrol agents. One Pythium species, Pythium insidiosum is also known to infect mammals.
  • The third group are the downy mildews, which are easily identifiable by the appearance of white "mildew" on leaf surfaces (although this group can be confused with the unrelated powdery mildews).

Classification of Oomycetes

Traditionally, this group was thought to include types of fungi, and, indeed, Fungi themselves were thought to be closely related to Plants. Many species of Oomycetes are still described and listed as types of Fungi.


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Scientific classification or biological classification is a method by which biologists group and categorize species of organisms. Scientific classification also can be called scientific taxonomy, but should be distinguished from folk taxonomy, which lacks scientific basis.
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Chromalveolata
Cavalier-Smith, 1998

Phyla
Heterokontophyta
Haptophyta
Cryptophyta
Alveolata
  • Ciliophora
  • Apicomplexa
  • Dinoflagellata


Chromalveolata
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Heterokontophyta

Typical classes

Colored groups
Chrysophyceae (golden algae)
Synurophyceae
Actinochrysophyceae (axodines)
Pelagophyceae
Phaeothamniophyceae
Bacillariophyceae (diatoms)
Bolidophyceae
Raphidophyceae
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Leptomitales

Scientific classification

Domain: Eukaryota

(unranked) Chromista

Phylum: Heterokontophyta

Class: Oomycetes

Order:
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Peronosporales are an order of water moulds (Class Oomycetes) which can be pathogenic.

Many diseases of plants are sometimes classified under this order, but are sometimes considered members of order Pythiales.
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Saprolegniales

Families

See text
Saprolegniales is an order of freshwater mould that includes the Saprolegniaceae and other families.
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American and British English spelling differences are one aspect of American and British English differences.

In the early 18th century, English spelling was not standardised. Different standards became noticeable after the publishing of influential dictionaries.
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Eukarya
Whittaker & Margulis, 1978

Kingdom: Protista*
Haeckel, 1866

Typical phyla
  • Chromalveolata
  • Chromista

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Eukarya
Whittaker & Margulis, 1978
(unranked) Opisthokonta

Kingdom: Fungi
(L., 1753) R.T. Moore, 1980[1]

Subkingdom/Phyla

Chytridiomycota
Blastocladiomycota

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Mycelium is the vegetative part of a fungus, consisting of a mass of branching, thread-like hyphae. The mass of hyphae is sometimes called shiro, especially within the fairy ring fungi. Fungal colonies composed of mycelia are found in soil and on or in many other substrates.
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Thallus, from Latinized Greek θαλλος (thallos), meaning a green shoot or twig, is an undifferentiated vegetative tissue of some non-mobile organisms, which were previously known as the thallophytes.
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cell wall is a fairly rigid layer surrounding a cell, located external to the cell membrane, which provides the cell with structural support, protection, and acts as a filtering mechanism. The cell wall also prevents over-expansion when water enters the cell.
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Cellulose is an organic compound with the formula (C6H10O5)n. It is a structural polysaccharide derived from beta-glucose.[1][2] Cellulose is the primary structural component of green plants.
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Chitin (C8H13O5N)n (IPA: [ˈkaɪtn̩]) is a long-chain polymer of beta-glucose that forms a hard, semitransparent material found throughout the natural world.
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septum (Latin: something that encloses; plural Septa) is a partition separating two cavities or spaces. Examples include:
  • Nasal septum: the cartilage wall separating the nostrils of the human nose.

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Phaeophyceae
Kjellman

The Phaeophyceae or brown algae, (singular: alga) is a large group of mostly marine multicellular algae, including many seaweeds of colder Northern Hemisphere waters.
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Bacillariophyceae

Orders
  • Centrales
  • Pennales
Diatoms (Greek: διά (dia) = "through" + τέμνειν
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Heterokontophyta

Typical classes

Colored groups
Chrysophyceae (golden algae)
Synurophyceae
Actinochrysophyceae (axodines)
Pelagophyceae
Phaeothamniophyceae
Bacillariophyceae (diatoms)
Bolidophyceae
Raphidophyceae
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A flagellum (plural: flagella) is a long, slender projection from the cell body, composed of microtubules and surrounded by the plasma membrane.
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spore is a reproductive structure that is adapted for dispersion and surviving for extended periods of time in unfavorable conditions. Spores form part of the life cycles of many plants, algae, fungi and some protozoans.
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A zoospore is a motile asexual spore utilizing a flagellum for locomotion. Also called a swarm spore, these spores are created by some algae and fungi to propagate themselves.
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An oospore is a thick-walled sexual spore that develops from a fertilized oosphere in some algae and fungi.
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''Plant pathology redirects here. For the journal, see Plant Pathology (journal).

Phytopathology (plant pathology) is the scientific study of plant diseases caused by pathogens (infectious diseases) and environmental conditions (physiological factors).
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Phytophthora

Species
Phytophthora arecae
Phytophthora botryosa
Phytophthora cactorum
Phytophthora cajani
Phytophthora cambivora
Phytophthora capsici
Phytophthora cinnamomi
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P. cinnamomi

Binomial name
Phytophthora cinnamomi
Rands

Phytophthora cinnamomi
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P. infestans

Binomial name
Phytophthora infestans
(Mont.) de Bary

Phytophthora infestans is an oomycete that causes the serious potato disease known as late blight or
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P. ramorum

Binomial name
Phytophthora ramorum
Werres et al. 2001

Sudden Oak Death (Phytophthora ramorum
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C. dentata

Binomial name
Castanea dentata
(Marsh.) Borkh.



The American Chestnut (Castanea dentata
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Pythium

Species

Pythium acanthicum
Pythium acanthophoron
Pythium acrogynum
Pythium adhaerens
Pythium amasculinum
Pythium anandrum
Pythium angustatum

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