Information about Planidium

A planidium is a specialized type of first-instar insect larva, seen in groups that are parasitoids; they are generally flattened, highly sclerotized, have legs, are quite mobile, and sometimes have eyes. They occur in the orders Hymenoptera, Coleoptera, Strepsiptera, and Diptera; specifically the beetle families Meloidae and Ripiphoridae, the fly family Acroceridae, and the parasitic wasp families Eucharitidae and Perilampidae. All Strepsiptera have planidial larvae. The first instar larva in Meloidae has three claws on each foot, and is therefore called a triungulin, but otherwise they are typical planidia. The term "triungulin" is sometimes used to refer to other planidial beetle or Strepsipteran larvae.

These larvae either wait for a passing host, or actively seek one out. In most cases they are phoretic, and ride on the adult form of the host or an intermediate vector, in order to gain access to the actual life stage attacked (typically, they enter the body of the host larva). In an unusual case, planidium larvae of beetles of the genus Meloe will form a group and produce a pheromone that mimics the sex attractant of its host bee species; when the male bee arrives and attempts to mate with the mass of larvae, they climb onto his abdomen, and from there transfer to a female bee, and from there to the bee nest to parasitize the bee larvae.[1] It is common for planidia to molt shortly after entering the host body, but they often postpone further development while the larva grows. This and subsequent molts usually involve a loss of legs and eyes, as well as de-sclerotization, a process called hypermetamorphosis.
instar is a developmental stage of arthropods, such as insects, between each moult (ecdysis), until sexual maturity is reached. Arthropods must shed the exoskeleton in order to grow or assume a new form.
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Insecta
Linnaeus, 1758

Orders
Subclass Apterygota
* Archaeognatha (bristletails)
* Thysanura (silverfish)
Subclass Pterygota
* Infraclass Paleoptera (Probably paraphyletic)

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larva (Latin; plural larvae) is a juvenile form of animal with indirect development, undergoing metamorphosis (for example, insects or amphibians).

The larva can look completely different from the adult form, for example, a caterpillar differs from a butterfly.
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parasitoid is an organism that spends a significant portion of its life history attached to or within a single host organism which it ultimately kills (and often consumes) in the process. Thus they are similar to typical parasites except in the certain fate of the host.
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Sclerosis or sclerotization is a hardening of tissue

In medicine

  • multiple sclerosis, a central nervous system disease which affects coordination
  • hippocampal sclerosis, a brain damage often seen in individuals with temporal lobe epilepsy

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Hymenoptera
Linnaeus, 1758

Suborders

Apocrita
Symphyta

Hymenoptera is one of the larger orders of insects, comprising the sawflies, wasps, bees, and ants.
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Strepsiptera
Kirby, 1813

Families

Mengenillidae
Mengeidae
Stylopidae
Bohartillidae
Corioxenidae
Halictophagidae
Callipharixenidae
Elenchidae
Myrmecolacidae

The Strepsiptera (occasionally known as
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Diptera
Linnaeus, 1758

Suborders

Nematocera (includes Eudiptera)
Brachycera

True flies are insects of the Order Diptera (Greek: di = two, and pteron
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Meloidae
Gyllenhal, 1810

Genera

See text.

Beetles in the Coleoptera family Meloidae are commonly known as blister beetles. There are approximately 2,500 known species worldwide.
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Ripiphoridae
Gemminger & Harold, 1870

Subfamilies

Hemirhipidiinae
Micholaeminae
Pelecotominae
Ptilophorinae
Ripidiinae
Ripiphorinae

The family Ripiphoridae (formerly spelled Rhipiphoridae
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Diptera
Linnaeus, 1758

Suborders

Nematocera (includes Eudiptera)
Brachycera

True flies are insects of the Order Diptera (Greek: di = two, and pteron
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Acroceridae

Subfamilies

Panopinae
Philopotinae
Acrocerinae
Acroceridae is a small family of odd looking flies most closely related to Nemestrinidae. There are about 520 species in 50 genera.
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parasitic wasp refers to a large, artificial assemblage of Hymenopteran superfamilies which are primarily parasitoids of other animals, mostly other arthropods. Many of them, such as the family Braconidae, are considered beneficial because they control populations of agricultural
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Eucharitidae
Latreille, 1809

Diversity
2 subfamilies
53 genera
c. 470 species

Subfamilies

Eucharitinae
Oraseminae

The Eucharitidae are a specialized group of ant parasitoids within the Chalcidoidea.
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Perilampidae
Latreille, 1809

Diversity
3 subfamilies
15 genera
c. 270 species

Subfamilies

Chrysolampinae
Perilampinae
Philomidinae

The Perilampidae
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Commensalism is a term employed in ecology to describe a relationship between two living organisms where one benefits and the other is not significantly harmed or helped. It is derived from the English word commensal
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pheromone is a chemical that triggers a natural behavioral response in another member of the same species. There are alarm pheromones, food trail pheromones, sex pheromones, and many others that affect behavior or physiology.
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Apidae

Subfamilies
  • Apinae
Honey bees, bumblebees, stingless bees, orchid bees and others
  • Nomadinae
Cuckoo bees
  • Xylocopinae
Carpenter bees

The Apidae
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Hypermetamorphosis is a kind of complete metamorphosis in which the different larval instars represent two or more different forms of larva. As the larva molts its morphology can change from that of a campodeiform larva to scarabaeiform (grublike) or to vermiform (maggotlike).
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