Information about Cedar Mountain Formation

The Cedar Mountain Formation is the name given to distinctive sedimentary rocks in eastern Utah that occur between the underlying Morrison Formation and overlying Naturita Formation (sometimes incorrectly called the Dakota Formation). It is composed of non-marine sediments, that is, sediments deposited in rivers, lakes and on flood plains. Based on various fossils and radiometric dates, the Cedar Mountain Formation was deposited during the last half of the Early Cretaceous, about 127 - 98 million years ago (mya). Dinosaurs occur throughout the formation, but their study has only occurred since the early 1990s. The dinosaurs in the lower part of the formation differ from those in the upper part. These two dinosaur assemblages, characterized by distinct dinosaurs, show the replacement of older, European-like dinosaurs with younger, Asian-like dinosaurs as the North American Continental Plate drifted westward. A middle dinosaur assemblage may be present, but the fossil record is not clear. The formation was named for Cedar Mountain in northern Emery County, Utah, where William Stokes first studied the exposures in 1944. Only recently did the 125 m (410 ft) thick formation get subdivided into smaller, distinctive beds called members. There is a debate as to whether there are five members (e.g., Kirkland et al. 1997) or four (e.g., Roca-Argemi and Nadon 2003) depending whether the Buckhorn Conglomerate is considered to be at the top of the Morrison Formation or at the base of the Cedar Mountain Formation; most geologists and paleontologist consider it part of the Cedar Mountain Formation. In ascending order the remaining members are the Yellow Cat Member, Poison Strip Sandstone, Ruby Ranch Member, and the Mussentuchit Member. Each of these members are named after a geographic area where they were first studied.

Stratigraphy

The Cedar Mountain Formation is sandwiched between the Morrison Formation below and the Dakota Formation and Mancos Shale above. The youngest date for Morrison just below the Cedar Mountain Formation is 148.1 + 0.5 Ma (Kowallis et al. 1998) or lower Tithonian. Typically, the Jurassic-Cretaceous boundary in western North America is marked by an unconformity of variable length, and typically signifies 10-49 million years of missing geologic time (Kowallis et al. 1998). This boundary between the Morrison and Cedar Mountain is commonly marked by a horizon of carbonate nodules (Stokes 1952; Aubrey 1998) or by highly polished pebbles that are allegedly gastroliths.

• The Buckhorn Conglomerate is considered the lowermost member of the Cedar Mountain Formation in the region of the San Raphael Swell by Stokes (1952). It is named for exposures near Buckhorn Reservoir near Cedar Mountain. Its position immediately below the Ruby Ranch Member suggests that it may be equivalent to the channel sandstones in the Yellow Cat Member and the Poison Strip Sandstone farther to the east. This idea is strengthened by the similar composition of the gravels in these members, but a direct correlation has not yet been established.
• The Yellow Cat Member is named for exposures near the Yellow Cat mining area north of Arches National Monument. It is limited to the eastern portions of the formation and is thickest near Arches National Monument. The member is composed of drab greyish mudstones and some lenses of sandstone. The mudstones were deposited on flood plains, and show evidence of ancient soil development called paleosols. The mudstones originated as flood deposits from river channels that are marked by the sandstone lenses. A recent radiometric date of 126 + 2.5 Ma places Yellow Cat Member in the Barremian (Kirkland and Madsen 2007), which verifies previous estimates based on fossil evidence (Kirkland et al. 1997).
• The Poison Strip Sandstone was named for prominent, cliff-forming sandstones in the Poison Strip uranium district north of Arches National Monument. It is actually a series of sandstones that were deposited in river channels, and lesser amounts of mudstones and limestones that were deposited on the flood plain and small ponds. Based on the position of the Poison Strip between the Yellow Cat and Ruby Ranch members, it probably was latest Barremian to earliest Aptian.
• The Ruby Ranch Member is the most widespread and distinctive member of the Cedar Mountain. It was named for exposures on the Ruby Ranch located southeast of Green River, Utah. The member is composed of maroon mudstones with irregular spheres of carbonate nodules. The nodules formed in ancient soils that developed in the mud deposited on the flood plain in a strongly seasonal, semiarid climate. Evaporation of groundwater during the dry season concentrated calcium carbonate and other minerals in the upper parts of the soil horizon. Radiometric dates place the upper portions of the Ruby Ranch in the late Aptian. Exhumed river channels in the Ruby Ranch indicate that stream flow during the Aptian was towards the northeast, the direction of the encrouching Western Interior Seaway.
• The Mussentuchit Member is the uppermost member of the Cedar Mountain Formation. It was named for exposures along Mussentuchit Wash southwest of the San Rafael Swell. It is composed predominately grey mudstones high in organic carbon from fossil plant material, as well as volcanic ash. The mudstones were originally deposited on a broad coastal plain with a high water table or with abundant rainfall. Thus, carbonate nodules are rare. A radiometric date of 98.37 + 0.07 Ma places the upper part of the member in the Lower Cenomanian.
• Although not part of the Cedar Mountain Formation, the Naturita Formation immediately overlies the Cedar Mountain and marks the encroaching Western Interior Seaway. The Naturita is not uniformly distributed and was eroded away in places by the advancing Seaway so that the marine shales of the Mancos Formation lay directly on the Mussentuchit or its equivalent. The name Dakota Formation has been improperly used for these strata.

Dinosaurs

The Cedar Mountain Formation is one of the last major dinosaur-bearing formations to be studied in the United States. Although sporadic bone fragments were known prior to 1990, serious research did not begin until that year. Since then, several organizations have conducted field work collecting dinosaurs, chiefly the Oklahoma Museum of Natural History, the Denver Museum of Nature & Science, the College of Eastern Utah, the Utah Geological Survey, Brigham Young University, and Dinosaur National Monument staff. This research indicates that at least two, possibly three dinosaur assemblages are contained within the formation.



The oldest of these assemblages is from the Yellow Cat, Poison Strip and basal Ruby Ranch members. The small, Ornitholestes-like theropod Nedcolbertia and the brachiosaurid sauropod Cedarosaurus may be considered as relics, with their closest relatives in the Morrison Formation. In contrast, the polacanthid ankylosaur Gastonia and a yet unnamed iguanodontid are similar to related forms from the Lower Cretaceous of southern England. These dinosaurs show that the connection between North America and Europe still existed during the Barremian. All of this changes, however, with the upper dinosaur assemblage from the top of the Ruby Ranch and Mussentuchit members. This upper assemblage shows greater similarities with Asian dinosaur assemblages from the same time. For example, the primitive ankylosaurid Cedarpelta is related to Gobisaurus and Shamosaurus from Mongolia, but is more primitive than either because it has teeth in the premaxilla. The upper assemblage also has a tyrannosaurid, a ceratopsian, and a pachycephalosaurs. Although not a dinosaur, the primitive mammal Gobiconodon is known from both Mongolia and the Mussentuchit Member. Evidence for a middle dinosaur assemblage between the older and younger ones is controversial because the evidence mostly depends on a single specimen of the ornithopod Tenontosaurus from high in the Ruby Ranch Member and the sauropod Astrodon from low in the Ruby Ranch. Regardless, the upper and lower dinosaur assemblages in the Cedar Mountain Formation document the separation of North America and Europe, the westward drift of North America, and its connection with Asia 10 to 15 million years later (Carpenter 2006).

Other Fossils

Besides dinosaurs, the Cedar Mountain Formation has produced a wealth of small fossils (a.k.a. microfossils), mostly teeth from a variety of vertebrates. Most of these specimens have been found in the Mussentuchit Member where they are collected by washing the rock through fine window screen. The teeth and other small fossils are picked from the residue (e.g., Cifelli et al. 1999).

• Fish include primitive fresh or brackish water sharks (e.g., Hybodus) and rays (c.f., Ischyrhiza), the lung fish (Ceratodus) and several bony fishes known from vertebrae. Lung fish are able to breathe air when pond water become poorly oxygenated, such as during the dry season.
• Amphibians include both salamanders (e.g. Albanerpeton) and frogs, but neither is common.
• Reptiles are more abundant and better studied. These include aquatic turtles (Glyptops, Naomichelys), at least one type of snake (Coniophis), and several different lizards, including teiids (Bicuspidon), a possible skinks and some extinct families (e.g., Paramacellodidae). Crocodiles are also present but their remains are fragmentary. They include Bernissartia, an unnamed atoposaurid, and unnamed pholidosaurid. At least one fragment of a large pterosaur is known from the base of the Mussentuchut Member. Unfortunately, it is too incomplete to identify to family or genus.
• Bird remains are very fragmentary because of their delicate structure. At least one aquatic bird is known. Based on the diversity of birds from the Early Cretaceous of China, other birds were probably present in Utah at this time as well.
• Mammals are the most thoroughly studied thanks to the work of Jeffrey Eaton and Richard Cifelli (summarized in Cifelli et al., 1999). They include triconodonts (e.g., Astroconodon), which have the molar cusps arranged in a single row; symmetrodonts (e.g., Spalacolestes; Spalacotheridium), characterized by molars having three cusps arranged in a triangle; multituberculates (e.g., Janumys; Cedaromys; Paxacimexomys), with their multiple rows of cusps on the molars; one of the earliest marsupials (Kokopellia), and several unnamed tribotheres, characterized by molars having three cusps that are typically asymmetrically arranged.

The various vertebrates are listed by member in the list below.

Non-vertebrate fossils are more widely distributed in the Cedar Mountain Formation. These include the distinctive reproductive structures of fresh water algae that are called charophytes. Charophytes are so distinctive that they are used to correlate strata of similar age, and thus were used to show that the Yellow Cat Member was time equivalent to Barremian age strata in England (Kirkland et al.1997). Ostracods, small crustaceans with clam-like shells, also occur in fresh water deposits, along with “finger-clams” or chonchostracans. Pollen have been found in the Mussentuchit Member and are important for reconstructing the environment. In a few places, large petrified logs are known, especially from the Poison Strip. These conifer logs are over a meter in diameter and indicate the presence of trees over 30 m (100 feet). The distinct wood of the tree fern Tempskia is occasional found as well.

List of fossil vertebrates by member

Vertebrates of the Yellow Cat Member

Osteichthyes Dipnoi :Ceratodus new species Actinopterygia :cf. Semionotus? sp. Amiiformes indet, :genus and species indeterminant Reptilia Chelonia :cf. Glyptops sp. Rhynchocephalia :Toxolophosaurus sp. Crocodilia :genus and species indeterminate Dinosauria Theropoda :Therizinosauroid ::Falcarius utahensis

:Dromaeosauridae ::Utahraptor ostrommaysorum :Family unknown ::Nedcolbertia justinhofmanni Sauropoda :Brachiosauridae ::Cedarosaurus weiskophae :Titanosauridae ::new genus and species Ornithopoda :Iguanodontidae ::new genus and species :Hadrosauria? ::Cedrorestes crichtoni Ankylosauria :Polacanthidae ::Gastonia burgei

Vertebrates of the Poison Strip Member

Dinosauria Theropoda :Dromaeosauridae ::Utahraptor sp.

Sauropoda :Titanosauriform ::Venenosaurus dicrocei Ornithopoda :Iguanodontidae ::Planicoxa venenica

Vertebrates of the Ruby Ranch Member

(upper and lower faunas combined)

Chondrichthyes Hybodontoidea :Hybodus sp. Reptilia Crocodilia :genus and species indeterminate Dinosauria Theropoda :Family Dromaeosauridae ::Deinonychus sp. :Family Allosauridae ? ::new genus and species ::cf. Acrocanthosaurus sp.

Sauropoda :Family Brachiosauridae ::cf. Astrodon sp. Ornithopoda :Family Iguanodontidae ::Tenontosaurus sp. Ankylosauria :Family Polacanthidae ::Gastonia new species :Family Nodosauridae ::new genus and species

Vertebrates of the Mussentuchit Member

Chondrichthyes Hybodontiformes :Hybodontidae ::Hybodus sp. :Polyacrodontidae ::Polyacrodus parvidens ::Lissodus spp. Orectolobiforines :Orectolobidae ::new genus and species Rajiformes :Ischyrhiza sp. :Pseudohypolophus sp. Myliobatiformes :cf. Baibisha n. sp. Orectolobiformes :Cretorectolobus sp. Osteichthyes Neopterygii, indeterminant :genus and species indeterminant Lepisosteiformes :?Lepisosteidae :genus and species indeterminant Pycnodontiformes :?Pycnodontidae ::genus and species indeterminant Amiiformes indet, :genus and species indeterminant Dipnoi :Ceratodontidae ::Ceratodus sp. Lissamphibia Caudata Albanerpetontidae :Albanerpeton cf. A. nexuosus Anura :Family indeterminant ::genus and species indeterminant Reptilia Testudines :Baenidae ::genus and species indeterminant :Pleurostemidae ::Naomichelys sp. :Glyptopsidae ::Glyptops sp. Squamata :Teiidae ::Harmodontosaurus emeryensis :Polyglyphanodontinae ::Dicothodon morrensis :Scincoidea ::Dimekodontosaurus masdeni :Scincomorpha ::Bothriagenys mysterion :?Paramacellodidae ::Pseudosaurillius sp. :family indeterminant ::Primaderma nessovi Serpentes :Aniliidae ::Coniophis sp. Crocodilia :Bernissartiidae ::Bernissartia sp. :Goniopholididae ::cf. Dakotasuchus sp. ::Polydectes sp. :Atoposauridae ::genus and species indeterminant

:Teleosauridae ::Machimosaurus sp. :Pholidosauridae ::genus and species indeterminant Pterosauria :genus and species indeterminant Dinosauria Theropoda :Dromaeosaurinae ::genus and species indeterminant :Veloceraptorinae ::genus and species indeterminant :Troodontidae ::genus and species indeterminant :Therizinosauridae ::genus and species indeterminant :Family indet. ::cf. Richardoestesia sp. :Tyrannosauridae? ::genus and species indeterminant Sauropoda :?Brachiosauridae ::genus and species indeterminant Ornithopoda :Iguanodontoidea ::Eolambia carolijonesia :Hypsilophodontidae ::cf. Zephyrosaurus sp. Pachycephalosauria :Pachycephalosauridae ::genus and species indeterminant Ceratopsia :Neoceratopsia ::genus and species indeterminant Ankylosauria :Nodosauridae ::Animantarx ramaljonesi ::new genus and species Avialae :?Hesperomithiformes :genus and species indeterminant Mammalia Triconodonta :Triconodontidae ::Astroconodon delicatus ::Corviconodon utahensis ::Jugulator amplissimus Multitubeirculata :Suborder incertae sedis ::Ameribaatar zofiae ::Janumys erebos ::Bryceomys intermedius ::Paracimexomys robisoni ::Paracimexomys perplexus ::Cedaromys parvus ::Cedaromys bestia Symmetrodonta :Spalacotheriidae ::Spalacotheridium noblei ::Spalacolestes inconcinnus ::Spalacolestes cretulablatta Tribotheria :Picopsidae ::genus and species indeterminant Pappotheriidae ::new genus and species Marsupialia ? :Family indeterminant ::Kokopellia juddi

data from Carpenter (2006), Cifelli et al. (1999), Kirkland and Madsen (2007), and The Paleobiology Database

Conclusions

The Cedar Mountain Formation is proving to contain one of the richest and most diverse Early Cretaceous dinosaur faunas in the world. The discoveries to date have revealed that the origin of some of the later Cretaceous dinosaurs may lie in the Cedar Mountain, but further work is needed to understand the timing and effects the changing position of the North American Plate had on dinosaurian evolution. Also needed is a better understanding of the effects that the changing North American Plate had on the non-dinosaur vertebrates.

References

Aubrey, W.M. 1998. A newly discovered, widespread fluvial facies and uncomformity marking the Upper Jurassic/Lower Cretaceous boundary, Colorado Plateau. Modern Geology, v. 22, p. 209-233.

Carpenter, K., 2006, Assessing dinosaur faunal turnover in the Cedar Mountain Formation (Lower Cretaceous) of eastern Utah, USA. Ninth International Symposium on Mesozoic Terrestrial Ecosystems and Biota, Abstract and Proceedings Volume, p. 21-25.

Cifelli, R. L., Nydam, R. L., Gardner, J. D., Weil, A., Eaton, J. G., Kirkland, J. I., and Madsen, S., 1999, Medial Cretaceous vertebrates from the Cedar Mountain Formation, Emery County, Utah: the Mussentuchit Local Fauna, in, Gillette, D.,Vertebrate Paleontology in Utah. Utah Geological Survey Miscellaneous Publication 99-1, p.219-242.

Kirkland, J.I. and Madsen, S.K. 2007. The Lower Cretaceous Cedar Mountain Formation, eastern Utah: the view up an always interesting learning curve. Fieldtrip Guidebook, Geological Society of America, Rocky Mountain Section. 1-108 p.

Kirkland, J.I., Britt, B., Burge, D., Carpenter, K., Cifelli, R., DeCourten, F., Eaton, J., Hasiotis, S., and Lawton, T., 1997b, Lower to Middle Cretaceous dinosaur faunas of the Central Colorado Plateau: a key to understanding 35 million years of tectonics, sedimentology, evolution, and biogeography. Brigham Young University Geology Studies, v. 42, p. 69-103.

Kowallis, B. J., Christiansen, E. H., Deino, A. L., Peterson, F., Turner, C.E., Kunk, M. J., and Obradovich, J. D., 1998, The age of the Morrison Formation. Modern Geology, v. 22, p. 235-260.

Roca-Argemi, X. and Nadon, G. C. 2003. The Buckhorn Conglomerate as the upper member of the Morrison Formation: new evidence from the type section, Cedar Mountain, Utah. Geological Society of America, Rocky Mountain Section, 55th Annual Meeting, Paper 14-1.

Stokes, W. L. 1944, Morrison and related deposits in the Colorado Plateau. Geological Society of America Bulletin v. 55, p. 951-992.

Stokes, W.L. 1952. Lower Cretaceous in the Colorado Plateau. American Association of Petroleum Geologists v. 36: 1766-1776.

Additional References (External links)

Dinosaurs of the Cedar Mountain Formation presented by the Utah Geological Survey

[https://scientists.dmns.org/sites/kencarpenter/Cedar%20Mountain%20storage/Introduction.aspx?PageView=Shared The Cedar Mountain Dinosaur Project]

Utah’s Newly Recognized Dinosaur Record from the Early Cretaceous Cedar Mountain Formation